1950s

Dr James Riley, an honours graduate of Edinburgh University, had to give up a surgical career because of an affliction of his hands, and instead became a radiotherapist in Dundee. He had always been fascinated by past heroes of medical research, and set out to carve his own niche in the study of mast cells. In the early 1950s, no one knew what their function was other than the mast cell granules being the storehouses of heparin.